Throughout the breeding season the male has to ensure that the temperature inside the mound is maintained at about 33°C - 91°F. The male puts his head into the mound and it is thought that he uses his tongue to measure the temperature. He must then either add or remove sand from the mound, according to the temperature within it and the season.
In spring, much heat is given off by the rotting organic material and the temperature needs to be reduced. At dawn, the male rakes off the sand covering the litter, and after allowing enough heat to escape he refills the hole with cool sand.
In summer, the male needs to protect the eggs from the heat of the sun. He adds more and more sand to the mound, which absorbs the sun's warmth. Then, in the cool of the morning, he removes the sand and scatters it in the colder air. When the sand has cooled down he puts it back on the mound.
During the cooler temperatures of autumn, the mound needs to be warmed up. To do this the male digs almost all of the soil away in the morning, spreading it out to be warmed in the sun. Throughout the hottest part of the day, he places hot layers of sand on the mound, so that as the sun goes down the nest has been rebuilt and is warm for the night.